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Kenozersky National Park occupies the south-western part of Plesetsky District and the north-western part of Kargopolsky District of the Arkhangelsk Oblast, at the border with the Republic of Karelia. The northern part of the Park is centered at the Lake Kenozero, one of the biggest lakes of the region. The headquarters of the Park are located in the village of Vershinino, on the northern shore of the lake. The Lake Kenozero is the source of the Kena River, a major left tributary of the Onega. The upper course of the Kena is located in the park, as well as the Pocha River, the major tributary of the Lake Kenozero, its source, Lake Pochozero, and the lower course of the main tributary of the Lake Pochozero, the Undosha River.
The southern part contains Lake Lyokshmozero and the upper course of the Lyokshma River, a tributary of Lake Lacha. Lakes Lyokshmozero and Kenozero are separated by a number of lesser lakes, including Lake Naglimozero and Lake Vilno.
Some areas along the Karelian border drain into the basin of the Vodla, and eventually to the Baltic Sea. Thus, the park contains the portion of the divide between the basins of Atlantic and Arctic oceans.